Comparative anatomy

Academic Year 2026/2027 - Teacher: VENERA FERRITO

Expected Learning Outcomes

The teaching aims to provide an insight into the anatomical and morpho-functional organization of vertebrates through a comparative approach . Macro and microscopic characters of organs  will be analyzed within an evolutionary context. The theoretical description of each apparatus will be accompanied by the analysis of anatomical models  during practical laboratories. The students will then have to acquire detailed knowledges morpho-functional  of vertebrates using the basic concepts of comparative surveys as scientific methodology.

Expected results

 Knowledge and understanding – By the end of the course, students should demonstrate an understanding of the anatomical and morpho-functional organisation of the groups belonging to the subphylum of Vertebrates, from a comparative and evolutionary perspective. They should be familiar with the fundamental stages of embryonic development, from gametogenesis to organogenesis, in vertebrates. Identify the macroscopic and microscopic characteristics of the various organ systems. Interpret the morphological diversity of vertebrates within an appropriate phylogenetic context.

Ability to apply knowledge and understanding – By the end of the course, students should be able to use the methodological concepts of comparative analysis as a scientific tool to analyse animal anatomical structures; to correlate the concepts of descriptive anatomy and embryology in order to interpret the stages and mechanisms of vertebrate evolution; to apply knowledge of descriptive anatomy and organ function in the recognition and description of anatomical specimens and models observed during laboratory sessions. Apply models of circulation, respiration and osmoregulation to explain the adaptation of different taxa to specific environments (freshwater, marine or terrestrial environments).

Independent judgement – By the end of the course, students must demonstrate that they are able to independently assess the biological and ecological mechanisms that drive evolutionary and adaptive processes in animals; to interpret structural changes in organs as functional responses to selective pressures from the environment; and to form critical judgements on the relationships of homology and analogy between the anatomical components of different classes of vertebrates.

Communication skills – By the end of the course, students must be able to present the morpho-functional diversity of vertebrates within a phylogenetic context in a clear, rigorous and structured manner; to use the specific technical and scientific terminology of comparative anatomy and developmental biology appropriately; and to describe complex biological processes using appropriate scientific language.

Learning skills – By the end of the course, students should have developed the ability to independently consult and study advanced scientific texts on anatomy, as well as the capacity to keep their methodological knowledge up to date by integrating macroscopic anatomical knowledge with the findings of new research conducted in the fields of molecular embryology and phylogeny

Course Structure

Lectures and laboratories

Required Prerequisites

Basic knowledge of  Vertebrates systematic zoology

Attendance of Lessons

Mandatory attendance

Detailed Course Content

Introduction to the study of comparative anatomy. An overview of the systematics and phylogeny of vertebrates. Fundamental aspects of vertebrate embryogenesis. Structural units of vertebrates and their organisational plan. Integument: structure and function of the integument and its derivatives. Skeletal system: general principles of the vertebral column. Anatomy and development of the vertebrae. Organogenesis and anatomy of the neurocranium. Dermatocranium. Splanchnocranium. Appendicular skeleton. Circulatory system: organogenesis, morphology and evolution of the heart. Evolution of the aortic arch system. Simple circulation, incomplete double circulation and complete double circulation. Urinary system: organogenesis of the vertebrate urinary system. An overview of renal function. Structure of the kidney in anamniotes and amniotes. Nervous system: Organogenesis, anatomy and evolution of the central nervous system. Peripheral nervous system. Sense organs: Organogenesis and anatomy of the eye. Organogenesis and anatomy of the vestibulo-cochlear organ.

Textbook Information

 I - Stingo et al., Anatomia Comparata. Edi. Ermes

II - Liem et al., Anatomia comparata dei Vertebrati una visione funzionale ed evolutiva. EdiSES.

III - E. Giavini, E. Menegola, Manuale di Anatomia Comparata. EdiSES

IV - G.C.Kent, Anatomia comparata dei Vertebrati. Ed Piccin.

V - Materiale didattico fornito dal docente (quanto proiettato a lezione)

VI - Kardong K.V., Vertebrates- Comparative Anatomy, Function, Evolution. Ed. Mc Graw Hill

Course Planning

 SubjectsText References
1The ChordatesV - I Cap. 2
2The evolutionary history of vertebratesV - I Cap. 2
3General concepts of Embryology V - I Cap. 3
4The early stages of vertebrate developmentV - I (Cap. 3)
5Vertebrate body planV - I Cap. 1 
6Integument of primitive aquatic vertebrates V - I Cap. 4
7ectotermi terrestri (Anfibi e Rettili)V - I Cap. 4
8endotermi terrestri (Uccelli eMammiferi)V - I Cap. 4
9An overview of the skeletal systemV - I Cap.5
10Development of chondrocranium and splanchnocraniusmV - I (Cap.5)
11Evolution of splanchnocranium and origin of jawV - I (Cap.5)
12Type of jaw suspensionV - I (Cap.5)
13Evolution of chondrocraniumV - I (Cap.5)
14Evolution of dermatocraniumV - I (Cap.5)
15Embryonic development of vertebraeV - I (Cap.5)
16Embryonic  development of the cardiovascular systemV - I Cap. 7
17Evolution of heart chambersV - I Cap. 7
18Evolution of arterial vessels systemV - I Cap. 7
19Evolution of aortic archesV - I Cap. 7
20Evolution of venous vessels systemV - I Cap. 7
21Evolution of lymphatic systemV - I Cap. 7
22Blood circulation modelsV - I Cap. 7
23Overview of urinary systemV -I Cap.10
24Embryonic development of urinary systemV- I Cap. 10
25Nefron structureV- I Cap. 10
26Osmoregulation in fishesV- I Cap. 10
27Excretion of nitrogen metabolism productsV- I Cap. 10
28Overview of Nervous systemV - I Cap.14
29Embryonic development of nervous systemV - I Cap.14
30Overview on spinal cord, spinal nerves and evolutionV - I Cap.14
31Medulla oblongataV - I Cap.14
32CerebellumV - I Cap.14
33MidbrainV - I Cap.14
34DiencephalonV - I Cap.14
35TelencephalonV - I Cap.14
36Cranial nervesV - I Cap.14
37Eye embryonic developmentV - I Cap.14
38Inner ear embryonic development (vestibular and auditory systems)V - I Cap.14

Learning Assessment

Learning Assessment Procedures

Oral exam

In itinere text

Examples of frequently asked questions and / or exercises

Development of the eye

Double and incomplete circulation

Kidney types

Cerebellum

VERSIONE IN ITALIANO