PALEOECOLOGY AND ENVIRONMENTAL CONSERVATION
Academic Year 2025/2026 - Teacher: Emanuela DI MARTINOExpected Learning Outcomes
1. Knowledge and understanding
The student will acquire a solid understanding of the fundamental concepts of paleoecology, developing the ability to observe, analyze, and describe the paleontological content of sedimentary successions.
2. Applying knowledge and understanding
The student will be able to apply the acquired knowledge to interpret paleontological assemblages and their stratigraphic variations. They will be capable of using fossil organisms as indicators of environmental changes, including those occurring over short time scales, with potential applications in the field of environmental conservation and conservation paleobiology.
3. Making judgements
The student will develop the ability to critically interpret paleontological data and use them in paleoenvironmental reconstructions. Independent and in-depth exploration of the topics covered will be encouraged, through both multi- and interdisciplinary approaches.
4. Communication skills
Through active participation in lectures, practical exercises, and laboratory activities, the student will learn to use the technical language of the discipline and to communicate acquired knowledge with clarity, accuracy, and scientific rigor. They will also be able to effectively present and discuss case studies, demonstrating skills in synthesis, argumentation, and the use of visual aids and bibliographic sources.
5. Learning skills
The student will develop an integrated and in-depth view of paleoecology and its relevance to environmental conservation and conservation paleobiology, enhancing their ability to independently expand their competencies through the study of recommended bibliographic resources, including reference texts and recent scientific publications. They will be able to update and deepen their knowledge critically, also in preparation for further studies or research activities.
Course Structure
· Lectures supported by PowerPoint presentations and videos.
· Laboratory activities aimed at learning techniques for processing paleontological and paleoecological samples, including the observation and analysis of fossils and fossiliferous rocks, also using microscopes, to identify fossil assemblages of paleoecological and paleoenvironmental interest. During the laboratory sessions, students will also learn methods for collecting, processing, and presenting quantitative data, which are essential for paleoecological and paleoenvironmental reconstructions, as well as for applied studies in environmental conservation and conservation paleobiology. Additionally, students will be trained in reading and understanding scientific publications and in presenting them seminar-style through PowerPoint presentations.
· Seminars delivered by researchers from our University or from other Italian and international research institutions, experts in specific topics and, in some cases, authors of the case studies discussed during the lectures.
· A possible field excursion to be held at the end of the course or at another date agreed upon with the students, for direct observation of fossil assemblages and paleobiocoenoses in the field.
To ensure equal opportunities and comply with current legislation, students with disabilities and/or specific learning disorders (DSA) can request a personal meeting to arrange any necessary compensatory measures, based on the educational objectives and individual needs. Students can also contact the CInAP (Centre for Active and Participated Integration – Disability and/or DSA Services) representative of our Department, Prof. Giorgio De Guidi.
Required Prerequisites
A basic knowledge of present-day physical geography (including the distribution of continents, oceans, and major seas), plate tectonics, and the general paleogeography of major continental masses and oceans from the Late Paleozoic onward is required. In addition, students should be familiar with the fundamental concepts of paleontology and stratigraphy.
Attendance of Lessons
Attendance to both lectures and laboratory sessions for at least 70% of the total scheduled hours is mandatory, in accordance with the teaching regulations. Attendance will be recorded by the course lecturer.
Detailed Course Content
Paleoecology and Ecology. Relationships between paleoecology and ecology: analogies and differences. Research approaches in both fields. Main challenges in paleoecological studies. Subdivisions of paleoecology and main operational units.
Marine Paleoecology. Life modes and trophic strategies of marine organisms. Biotic and abiotic factors influencing organism distribution and their spatio-temporal variation, with a focus on benthic communities. Depositional environments and main physiographic units. Vertical zonation of the marine realm.
Relationships between benthic organisms and substrate. Hard, soft, ephemeral, and anthropogenic substrates: distribution and characteristics. Influence of substrate on organism distribution. Morphological adaptations related to substrate and habitat. Evolution of seafloor conditions.
Benthic bionomy and zonation. Zonation of the Benthic Domain: zones, systems, and major biocoenoses of geological, paleontological, and conservation interest. Heterogeneous assemblages as indicators of environmental instability (tectonic and/or climatic). From living associations (biocoenoses) to fossil assemblages (paleobiocoenoses): interpretative criteria and limitations.
Methods of study. Sampling techniques and analysis of fossil assemblages in the field and laboratory. Practical exercises: sample processing and microscope observations.
Environmental conservation and conservation paleobiology. Definition, aims, and potential applications. Use of historical and fossil data for the conservation of modern ecosystems and species. Concepts of ecological baseline and shifting baseline. Ecosystem variability in response to climate change and anthropogenic impact.
Case studies. Selected examples of studies focused on paleoecological and/or paleoenvironmental reconstructions, and on applications to environmental conservation and conservation paleobiology.
Textbook Information
Slides from all lectures and scientific publications provided by the teacher, to be supplemented with selected chapters from:
· AA. VV. 2020. Manuale di Paleontologia, Fondamenti-Applicazioni. Edizioni Idelson-Gnocchi.
· Benton M.J., Harper D.A. 2020. Introduction to Paleobiology and the Fossil Record. Wiley-Blackwell.
· Cognetti G., Sarà M., Magazzù G. 1999. Marine Biology. Calderoni, Bologna.
· Dietl G.P., Flessa K.W. 2017. Conservation Paleobiology: Science and Practice. University of Chicago Press.
· Kinne O. 1982. Marine Ecology. A Comprehensive, Integrated Treatise on Life in Oceans and Coastal Waters. John Wiley & Sons Inc.
· Margalef R. 1985. Key Environments. Western Mediterranean. Pergamon Press.
· Raffi S., Serpagli E. 1993. Introduzione alla Paleontologia. Edizioni Utet.
Further details are available in the "Course Planning" section.
Course Planning
| Subjects | Text References | |
|---|---|---|
| 1 | Paleoecology and Ecology: Definitions, similarities, and differences.Paleoecological issues: displacement and reworking.Uniformitarianism and Taxonomic Uniformism: caution in application.Subdivisions of Paleoecology: marine paleoecology, continental paleoecology, paleoecology of transitional environments.Different study Approaches in Ecology and Paleoecology: autoecology, synecology, population dynamics.Main operational units in Paleoecology: Individual, population, assemblage, biocoenosis, habitat, ecological niche, biotope, ecosystem. | Lecture slides; Raffi & Serpagli Chapter 6; AA VV. Manuale di Paleontologia, Fondamenti-Applicazioni Chapter 21; Benton & Harper Chapter 4 |
| 2 | Environmental and ecological factors: biotic and abiotic (physical and chemical) factors. Eurivalent, stenovalent, euriecic, and steniecic organisms. Seafloor morphology. Main depositional environments. Morphology and distribution of the oceans through geological time. Hypsographic curve. Methodologies for seafloor mapping. | Lecture slides; Cognetti et. al. Chapter 2; AA VV. Manuale di Paleontologia, Fondamenti-Applicazioni Chapter 21 |
| 3 | Modes of life of organisms: habitat and mobility. Plankton, benthos, and nekton.Overview of trophic relationships: trophic groups and food webs in the marine environment. Vertical zonation of the marine environment. Substrate types: hard and soft substrates. Hard and soft substrates in coastal areas, on the continental shelf, on the slope, and in the abyssal plain. Ephemeral substrates. Anthropogenic substrates. Characteristics of hard substrates influencing the distribution of organisms. Hard-substrate benthos. Characteristics of soft substrates influencing the distribution of organisms. Soft-substrate benthos. | Lecture slides; Raffi & Serpagli Chapter 6; Cognetti et al. Chapter 8 |
| 4 | Physical factors: depth, pressure, temperature, and light availability. Characteristics and distribution of organisms in relation to these environmental factors. | Lecture slides; Raffi & Serpagli Chapter 6; AA VV. Manuale di Paleontologia, Fondamenti-Applicazioni Chapter 21; Cognetti et al. Chapters 3 and 5; Margaleff Chapter 4 |
| 5 | Chemical factors: salinity, density, dissolved gases (oxygen and carbon dioxide), pH, and trace elements (nutrients). Characteristics and distribution of organisms in relation to these factors. | Lecture slides; Raffi & Serpagli Chapters 6 and 9; AA VV. Manuale di Paleontologia, Fondamenti-Applicazioni Chapter 21; Cognetti et al. Chapter 3; Margaleff Chapter 5 |
| 6 | Sea movements: wave motion, tsunami waves, tides, and currents. Characteristics and distribution of organisms in relation to sea movements. Challenges and adaptive strategies. Ecophenotypic variability. Tsunami deposits in the fossil record. Circulation patterns in the Mediterranean Sea. | Lecture slides; Raffi & Serpagli Chapters 6; AA VV. Manuale di Paleontologia, Fondamenti-Applicazioni Chapter 21; Cognetti et al. Chapter 4 |
| 7 | Biotic factors and biotic interactions: mobility (vagile, sedentary, and sessile organisms), trophic relationships, symbiotic interactions (mutualism, commensalism, parasitism), competition for space/substrate, epibiosis, bioimmuration, xenomorphosis. Post-mortem colonization of skeletons. | Lecture slides; Raffi & Serpagli Chapters 6; Cognetti et al. Chapter 7 |
| 8 | Benthic bionomy. Definition of zone and system. Vertical zonation of the marine environment. Abiotic factors that determine the boundaries of benthic zones. Zonation according to the Endoume School. Definition of (paleo)biocoenosis. Benthic zonation and major (paleo)biocoenoses of paleoecological interest in the Mediterranean. Categories of species within a (paleo)biocoenosis: exclusive characteristic species, preferential characteristic species, accompanying species, accidental species. | Lecture slides; Raffi & Serpagli Chapters 6; Cognetti et al. Chapter 9; Kinne Chapters 8 e 9; Margaleff Chapter 8 |
| 9 | Benthic (Paleo)biocoenosis of supralittoral, mesolittoral, and infralittoral zones | Lecture slides and supplementary handouts provided by the lecturer; Cognetti et al. Chapter 9 |
| 10 | (Paleo)Biocoenoses of the circalittoral and bathyal Zones. (Paleo)Biocoenoses independent of bathymetric zones. Heterogeneous assemblages. | Lecture slides and supplementary handouts provided by the lecturer; Cognetti et al. Chapter 9 |
| 11 | From living associations to fossil assemblages. Thanatocoenosis, taphocoenosis, orictocoenosis. Mixed residual fossil community. | Lecture slides; Raffi & Serpagli Chapters 6 |
| 12 | Sampling techniques and methods of analysis of fossil assemblages in the field and laboratory. | Lecture slides; AA VV. Manuale di Paleontologia, Fondamenti-Applicazioni Chapter 5 |
| 13 | Environmental conservation and conservation paleobiology. Definitions, aims, and potential applications. Use of historical and fossil data for the conservation of modern ecosystems and species. Concepts of ecological baseline and shifting baseline. Ecosystem variability in response to climate change and anthropogenic impact. | Lecture slides and supplementary handouts provided by the lecturer; Dietl & Flessa selected chapters 1, 5, 7, 8, 9, 10, 12, 13, 14 |
| 14 | Case studies: selected examples of studies focused on paleoecological and/or paleoenvironmental reconstructions, and on applications to environmental conservation and conservation paleobiology. | Scientific papers provided by the teacher |
Learning Assessment
Learning Assessment Procedures
Learning will be assessed through a final oral examination, scheduled according to the official calendar published on the degree program website. The questions will follow the format of those provided in the example list and will focus primarily on benthic bionomy, including relevant definitions and key concepts. During the exam, students will also be asked to present the characteristics of a biocoenosis of their choice, illustrating its relevant paleoecological aspects.
In addition, a midterm assessment is planned during the teaching break period established by the LM-74/79 degree program. This assessment will consist of an individual seminar-style presentation (PowerPoint format), in which the student will present one of the case studies discussed in class, based on a scientific publication selected by the teacher and agreed upon with the students. This assignment will assess the student’s ability to understand and communicate a paleoecological analysis, synthesize complex information, and argue with clarity and scientific rigor. It will also provide an opportunity to strengthen oral presentation skills and the ability to critically discuss case studies.
Examples of frequently asked questions and / or exercises
· Description of a paleoecologically relevant biocenosis of the student’s choice.
· What is meant by the term biocenosis?
· How is an exclusive characteristic species defined?
· What are accompanying species in a biocenosis?
· What are accidental species within a biocenosis?
· What are the main paleoecologically relevant biocenoses of the supralittoral zone?
· Which biocenoses characterize the mesolittoral zone and are relevant to paleoecology?
· What paleoecologically important biocenoses are found in the infralittoral zone?
· What are the typical biocenoses of the circalittoral zone relevant to paleoecology?
· Which biocenoses of paleoecological interest are found in the bathyal zone?
· What is meant by heterogeneous population and what is its paleoecological significance?
· How is a thanatocoenosis defined and what importance does it have in paleoecology and environmental conservation?
· What is a taphocoenosis and what is its significance?
· What is meant by orictocenosis?
· What is meant by residual fossil community?