PALEONTOLOGY WITH LABORATORY 1

Academic Year 2026/2027 - Teacher: MARIA ANTONIETTA ROSSO

Expected Learning Outcomes

Knowledge and understanding: Acquisition of the ability to observe, describe, and recognize different types of fossils. Knowledge of the most common fossilization processes for preserving evidence of past life. Knowledge of the evolutionary process and the history of life, from the emergence of the first organisms to the present-day. Acquisition of the concept of index fossils and the possibility of using fossils for dating rocks; ability in the identification of some stratigraphically important species. Knowledge of the effectiveness of using fossils for further purposes (mainly in geology) and ability to promote the fruition of fossil heritage.

Ability to master binomial nomenclature and understand the organization of systematics; ability to recognize fossils of the main marine invertebrate taxa by identifying their main characteristics in fossil samples and fossiliferous rocks; ability to recognize some species and high-rank taxa, primarily some of regional importance.

Ability to apply knowledge and understanding: develop the ability to search for and locate various types of fossils in the field, and to identify them using relevant literature, including online resource, thereby using them for dating purposes and the interpretation of paleoenvironments.

Students are also expected to demonstrate a high degree of autonomy in data collection and literature selection, both for the purpose of ongoing personal development (lifelong learning) and for communicating their ideas to others in a clear and engaging manner.

Course Structure

Teaching will be carried following the plan reported below. In the remote presence of restrictions, necessary changes will be made, according to.the University rules.

1. lectures with PP presentations, diagrams and, possibly, films,

2. laboratories with examination of fossils and fossiliferous rocks in order to learn to observe, describe and recognize the fossils, both isolated and within rocks. During laboratories, we will also learn methods for the collection, processing and presentation of simple data to describe individual taxa and for the documentation and enhancement of the fossil heritage. Formation about how to search online for scientific purposes, mostly if the informatic room will be available for laboratories.

3. seminars by researchers from Italian or foreign Universities, who are specialists of particular topics. Possibly at least one on Cenozoic mammifers.

4. field excursion at the end of the course.

To guarantee equal opportunities and in compliance with the laws in force, students with disabilities and/or SLDs can ask for a personal interview in order to plan any compensatory and/or dispensatory measures, based on the didactic objectives and specific needs.

It is also possible to contact the referent teacher CInAP (Center for Active and Participatory Integration - Services for Disabilities and/or SLD) of our Department, prof. Giorgio De Guidi.

At the request of students, topics of particular interest (or related topics) may be developed to carry out the final reports for the graduation.

Required Prerequisites

General knowledge about chemistry, mineralogy and general geology, especially in relation to sedimentary rocks and their sedimentation environments.

Basic knowledge of zoology.

Attendance of Lessons

Mandatory.

To achieve attendance, it is necessary to attend at least 70% of the lessons, as required for all disciplines in the teaching rules of the course in Geological Sciences.

Attendance will be monitored directly by the teacher.

Detailed Course Content

Definition of Paleontology: history, articulation and employ. Concept of fossil.

Paleontological systematic: concept of biological and palaeontological species, morphospecies, cronospecies, intraspecific variability. Nomenclature: the species of Linneous and the binomial nomenclature. The International Code. The classification of organisms. Parataxonomy.

Taphonomy: mineralised and unmineralised parts of organisms; death. Biostratinomic processes: necrolisis, bioerosion, dissolution, transport. Burial, incorporation in resins, incrustation. Fossilisation: organic matter; mineralised parts. Types of fossil-bearing and exceptional fossil deposits, oriented distributions of fossils and concentrations: interpretation and use. Lithogenetic relevance of some fossil groups.

Fossils in the field and in the lab: main methods for observation, sampling, analysis, cataloging and exploitation.

Fossil and evolution: information about evolution, microevolution, macroevolution. History of life on Earth.

Fossils and stratigraphy: fossils as a tool for dating rocks and make correlations.

Information on fossil traces: formation, conservation, classification and palaeontological interest.

Information on palaeoecology; the distribution of organisms as a key for palaeoenvironmental, palaeobiogeographic and palaegeographic reconstructions.

Global change: fossils as climatic indicators with examples from the Mediterranean history.

Fossils as cultural heritage: conservation and enjoyment.


Structure, organisation and classification, as well as rough information on autoecology, functional morfology and stratigraphic distribution of selected taxa selezionati from marin environments. Introduction to Cyanoficeae, Diatomeae, Radiolaria, Foraminifera, Coccolithophorida, Coralline algae; Tabulata,

Heliolithiida, Rugosa, Scleractinia, Octocorallia, Serpuloidea, Polyplacophora, Scaphopoda, Bivalvia, Gastropoda, Cephalopoda, Bryozoa, Brachiopoda, Crinoidea, Echinoidea, Trilobita, Ostracoda, Cirripedia.

Textbook Information

Texts chosen from those listed below, covering both general paleontology and paleontological systematics. The former is highly recommended among the Italian texts; Doyle 1996 among the English text.


Course Planning

 SubjectsText References
1Paleontology: origin and history; structure. The concept of fossil. Types of fossils: fossil bodies, models and impressions, traces.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
2Species. Biological and paleontological concepts. Binomial nomenclature. Introduction to systematics. Classification.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
3Possibility of fossilization of the different groups of organisms. Soft tissue and mineralized parts. Taphonomy. Death. Decomposition. Fossilization of organic matter. Exceptional fossilizations.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
4Chemical and mineralogical composition of skeletons and their preservation possibility. Stability of the minerals constituting the skeletons.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
5Transport and its consequences: fragmentation; abrasion. Selective transport. Oriented deposition. Use of the results of these processes in paleoenvironmental reconstructions.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
6Destructive biostratinomic processes: Disarticulation, Maceration, Bioerosion, Pre-diagenetic dissolution. Conservative biostratinomic processes: Encrustation.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
7Temporary and permanent burial. Exhumation and reworking: recognition and implications. Fossil diagenesis. Fossilization of mineralized parts. Diagenetic dissolution.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
8Internal moulds and external casts: how they form; significance and interest for paleontological taxonomy and geology. Deformations: types, meaning, and use for inferences.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
9Types of deposits and interpretation. Accumulations and concentrations of fossils. How they form and what they tell us. Fossils as constituents and builders of sedimentary rocks.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
10Introduction to ecology and paleoecology. Methods and objectives. Paleoecological, paleobiogeographic, and paleogeographic reconstructions.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
11Trace fossils: their formation and preservation. Stratinomic and ethological classification. Use of trace fossils in paleoenvironmental reconstructions. Bioturbation and sediment homogenization.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
12Fossils and the evidence of evolution. Historical and current evolutionary theories. Notes on the genetic basis of evolution.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
13Microevolution: The Origin of Species. Modes of Speciation.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
14Macroevolution: the origin of high-rank taxa. Basic and mass extinctions. Adaptive radiations.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
15Irreversibility of evolution and its consequences for stratigraphy. The concept of index fossils. Requirements and characteristics of index fossils. Relative dating and the geologic time table. Correlations. Examples of index fossils in the Sicilian area.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
16The history of life on Earth and some of its key stages. The evolution of the Mediterranean as documented by fossils.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
17Past climate variations and future prospects. An overview of the basic principles of conservation, cataloging, and display of fossil finds.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
18Morphological characteristics, classification, distribution and paleontological importance of the main groups of Protists.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
19Morphological characteristics, classification, distribution and paleontological importance of the main groups of coralline algae, archeocyatids and sponges.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
20Morphological characteristics, classification, distribution and paleontological importance of the main groups of conularids, tabulata, heliolithids and octocorals.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
21Morphological characteristics, classification, distribution and paleontological importance of the main groups of  tetracorals and  esacorals.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
22Morphological characteristics, classification, distribution and paleontological importance of the main groups of monoplacophorans, polyplacophorans,  gastropods and scaphopods.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
23Morphological characteristics, classification, distribution and paleontological importance of the main groups of bivalves.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
24Morphological characteristics, classification, distribution and paleontological importance of the main groups of cephalopods.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
25Morphological characteristics, classification, distribution and paleontological importance of the main groups of serpulids and bryozoans.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
26Morphological characteristics, classification, distribution and paleontological importance of the main groups of brachiopods.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
27Morphological characteristics, classification, distribution and paleontological importance of the main groups of echinoderms (echinoids and crinoids).Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
28Morphological characteristics, classification, distribution and paleontological importance of the main groups of trilobites, ostracods and cirripeds.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.
29Recognizing of some trace fossils. Recognizing of fossil bodies, models, fossil moulds and casts.Preferably AAVV edited by the Società Paleontologica Italiana. Manuale di Paleontologia. Fondamenti. Applicazioni. Idelson Gnocchi, 2020.

Learning Assessment

Learning Assessment Procedures

Learning will be verified through an oral examination at the end of the course according to the scheduled timetable appropriately advertised on the study course website.

Examination also includes identification of fossils and fossiliferous rocks and the possible solution of simple exercises.

An ongoing test is planned to be carried out on a date to be agreed with the attending students, but within the period specifically defined by the L34 Study Course for the suspension of teaching.

In the ongoing test, students will prove to be able to describe and recognize at least one fossil (provided by the teacher) and answer a few questions, some of which are multiple choice and others are short open-ended.

Examples of frequently asked questions and / or exercises

1. How many types of fossils do you know?

2. What are body fossils?

3. What are trace fossils? How are they produced and how are they preserved in the geological record?

4. What is taphonomy? How many and what taphonomic phases can be recognised?

5. What are biostratinomic processes?

6. What is a fossiliferous deposit? how many types of fossiliferous deposits do you know?

7. Stratigraphic distribution of species: examples from different eras.

8. What is macroevolution?

9. What is speciation?

11. What does pre-adaptation consist of?

12. What are mass extinctions? and which ones do you know?

13. Do you know fossils documenting key events in the history of the Mediterranean?

14. How would you organize a fossil exhibition?

15. What are the morphological characteristics used for the classification of irregular echinoids?

16. What are the morphological characteristics of bryozoans and what are the main characters used to distinguish the two classes we dealt with?

17. Similarities and differences between tetracorals and hexacorals.

18. What are the characteristics common to ammonoids and nautiloids and what characters are useful  to distinguish them?

19. Fossils characterized by the presence of two valves can be very distant taxonomically: which ones do you know and what are the differences?

20. What are the main characters we can use for  the distinction of the fossils of the unicellular organisms that we studied in the course?

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